The Scorpion Files - Diplocentridae

# Species: 2858

DIPLOCENTRIDAE
Karsch, 1880

This family consists of 10 genera and 140 species [Updated 24.07.23.23]. They are closly related to the Scorpionidae, but can be seperated from these by the presence of a subaculear spine on the telson (the Scorpionids lack this feature). Diplocentrids are located in North America (Mexico, Southwestern USA [Texas, New Mexico, and Arizona]), Central America (Belize, Costa Rica, El Salvador, Guatemala, Honduras, Nicaragua) and South America (Colombia, Venezuela), Caribbean (Greater & Lesser Antilles), Asia (Egypt, Iran, Israel, Jordan, Lebanon, Oman, Saudi Arabia, Syria, Yemen).

Scorpions in this family are occasionally kept in captivity (especially in the USA).

Diplocentrid scorpions are generally not medically important (but a few serious cases involving the genus Nebo in the Middle East have been reported.) See species file for more information.

Nebo photo by Dave Gaban (C).

SPECIES FILES:
This list of genera and species in The scorpion Files is mainly based on Fet et al.(2000). September 2019: Subspecies are included in the list. They included the family Diplocentridae as a subfamily in Scorpionidae. New studies in 2017-2019 and the opinion from several specialists support a family status for Diplocentridae and I have in July 2019 chosen to reinstate Diplocentridae as a family in The Scorpion Files. I try to update the list as additions and changes are published. * denotes changes after Fet et al. (2000). I will be grateful for information about new development in the taxonomy of this family. For information about synonyms and bibliographies, see Fet et al. (2000)

Bioculus Stahnke, 1968 (5) B. caboensis (Stahnke, 1968) B. cerralvensis Stahnke, 1968 B. comondae Stahnke, 1968 B. cruzensis Stahnke, 1968 B. parvulus Martin-Frias, 2004*	Cazierius Francke, 1978 (13) C. alayoni Armas, 1999* C. asper Teruel, 2006* C. cayacoa Teruel, Jimenez & de los Santos, 2021* C. chryseus Teruel & Armas, 2006* C. ciguayo Teruel, Jimenez & de los Santos, 2021* C. garridoi Armas, 2005* C. granulosus Teruel, 2013* C. gundlachii (Karsch, 1880) C. neibae Kovarik & Teruel, 2014* C. paradoxus Teruel & Diaz, 2004* C. parvus Armas, 1984* C. politus (Pocock, 1898) C. torrei (Moreno, 1938)*
Cryptoiclus Teruel & Kovarik, 2012 (1) C. rodriguezi Teruel & Kovarik, 2012	Didymocentrus Kraepelin, 1905 (12) D. armasi Teruel & Rodriguez, 2008* D. hasethi (Kraepelin, 1896) D. hummelincki Francke, 1978 D. jaumei Armas, 1976* D. krausi Francke, 1978 D. lesueurii (Gervais, 1844) D. martinicae Teruel & Questel, 2020* D. minor Francke, 1978 D. nitidus (Hirst, 1907) D. sanfelipensis Armas, 1976* D. trinitarius (Franganillo, 1930) D. waeringi Francke, 1978
Diplocentrus Peters, 1861 (65) D. actun Armas & Palacios-Vargas, 2002* D. anophthalmus Francke, 1977 D. bellator Teruel, 2003* D. bereai Armas & Martin-Frias, 2004* D. bicolor Contreras-Felix & Santibanez-Lopez, 2011* D. chol Francke, 2007* D. churumuco Francke & Ponce Saavedra, 2005* D. chiapasensis Beutelspacher & Armas, 1998* D. coddingtoni Stockwell, 1988 D. colwelli Sissom, 1986 D. coylei Fritts & Sissom, 1996 D. cozumel Beutelspacher & Armas, 1998* D. cueva Francke, 1978 D. diablo Stockwell & Nilsson, 1987 D. duende Santibanez-Lopez & Gonzalez-Santillan, 2017* D. ferrugineus Fritts & Sissom, 1996 D. formosus Armas & Martin-Frias, 2003* D. franckei Santibanez-Lopez, 2014* D. gertschi Sissom & Walker, 1992 D. gladiator Beutelspacher & Trujillo, 1999* D. hoffmanni Francke, 1977 D. insularis Sagastume-Espinoza, Longhorn & Santibanez-Lopez, 2015* D. izabal Armas & Trujillo, 2016* D. jaca Armas & Martin-Frias, 2000* D. keyserlingii Karsch, 1880 D. kraepelini Santibanez-Lopez, Francke & Prendini, 2013* D. lachua Armas, Trujillo & Agreda, 2011* D. landelinoi Trujillo & Armas, 2012* D. leptomanus Villa-Corella, Silva-Kurumiya, Barrales-Alcala, Van Devender & Francke, 2023* D. lindo Stockwell & Baldwin, 2001* D. longimanus Santibanez-Lopez, Francke & Athanasiadis, 2011* D. lourencoi Stockwell, 1988 D. lucidus Stockwell, 1988 D. luisae Guijosa, 1973 D. majahuensis Baldazo Monsivaiz, 2003* D. motagua Armas & Trujillo, 2009* D. maya Francke, 1977 D. melici Armas, Martin-Frias & Berea, 2004* D. mexicanus Peters, 1861 D. mexicanus mexicanus Peters, 1861 D. mexicanus karschi Sissom & Francke, 1998 D. mitchelli Francke, 1977 D. mitlae Francke, 1977 D. montecristo Armas & Martin-Frias. 2000* D. ochoterenai Hoffmann, 1931 D. ornatus Stockwell, 1988 D. oxlajujbaktun Trujillo & Armas, 2012* D. peloncillensis Francke, 1975 D. perezi Sissom, 1991 D. popti Trujillo & Armas, 2016* D. rectimanus Pocock, 1898 D. reddelli Francke, 1977 D. roo Armas & Martin-Frias, 2005* D. sagittipalpus Santibanez-Lopez, Francke & Prendini, 2013* D. santiagoi Stockwell, 1988 D. silanesi Armas & Martin-Frias. 2000* D. sinaan Armas & Martin-Frias. 2000* D. sissomi Santibanez-Lopez, Francke & Prendini, 2013* D. spitzeri Stahnke, 1970 D. steeleae Stockwell, 1988 D. taibeli (Caporiacco, 1938) D. tehuacanus Hoffmann, 1931 D. tehuano Francke, 1977 D. tenango Santibanez-Lopez & Francke, 2008* D. whitei (Gervais, 1844) D. williamsi Sissom & Wheeler, 1995 D. zacatecanus Hoffmann, 1931	Heteronebo Pocock, 1899 (19) H. barahonae Teruel, de Armas & Kovarik, 2015* H. bermudezi (Moreno, 1938) H. caymanensis Francke, 1978 H. cicero Armas & Marcano-Fondeur, 1987 H. clareae Armas, 2001* H. dominicus Armas, 1981 H. elegans Francke, 1978 H. forbesii Pocock, 1899 H. granti Pocock, 1899 H. jamaicae Francke, 1978 H. monticola (Armas, 1999)* H. morenoi (Armas, 1973)* H. nibujon Armas, 1984 H. oviedo (Armas, 1999)* H. portoricensis Francke, 1978 H. pumilus Armas, 1981 H. scaber (Pocock, 1893)* H. vachoni Francke, 1978 H. yntemai Francke & Sissom, 1980
Kolotl Santibanez-Lopez, Francke & Prendini, 2014* (2) K. magnus (Beutelspacher & Lopez-Forment, 1991)* K. poncei (Francke & Quijano-Ravell, 2009)*	Nebo Simon, 1878 (10) N. flavipes Simon, 1882 N. franckei Vachon, 1980 N. grandis Francke, 1980 N. henjamicus Francke, 1980 N. hierichonticus (Simon, 1872) N. jazanensis Abu Afifeh, Aloufi, Al-Saraireh & Amr, 2023* N. omanensis Francke, 1980 N. poggesii Sissom, 1994 N. whitei Vachon, 1980 N. yemenensis Francke, 1980
Oiclus Simon, 1880 (7) O. ardens Ythier, 2019* O. cousteaui Ythier, 2019* O. nanus Teruel & Chazal, 2010* O. purvesii (Becker, 1880) O. purvesii purvesii (Becker, 1880) O. purvesii sabae Francke, 1978 O. questeli Teruel, 2008* O. tipunch Ythier, 2019* O. tite Ythier, Jourdan & Malglaive, 2022*	Tarsoporosus Francke, 1978 (6) T. anchicaya Lourenço & Florez, 1990 T. flavus (Gonzalez-Sponga, 1984) T. kugleri (Schenkel, 1932) T. macuira Teruel & Roncallo, 2007* T. tizopedrosoi Bedoya-Roqueme & Lira, 2022* T. yustizi (Gonzalez-Sponga, 1984)