With 73 genera (one extinct) and 529 species (until 1998),
this family is the largest of the scorpion families. This
family is widespread around the world (not found in Antarctica
and New Zealand), and are found in tropical, subtropical and
partly in temperate habitats.
The members of this family are small and medium sized
scorpions with a triangular sternum (some genera have a more
pentagonal sternum). In many genera (e.g., Androctonus,
Apistobuthus, Parabuthus, and other Old World
genera) the cauda is usually strong and powerful and the
pedipalps are often very slender whereas other genera have
elongate appendages, especially the males (e.g.,
Centruroides, Lychas, Isometrus and
related genera). Many species are yellow or brown (or
variations of these colors), but black forms are also present.
Some species have quite intricate and attractive patterns
(e.g., Lychas and Isometrus) and colours (e.g.,
Centruroides and Uroplectes). Sizes range from
about 20 mm (like Microtityus and Microbuthus)
to over 120 mm (as in Androctonus, some
Centruroides, Apistobuthus and others).
Several species in this family are highly toxic, but fewer
than 20 can be lethal to man. The most notorious species are
found in the genus Androctonus, Centruroides, Hottentotta,
Leiurus, Parabuthus and Tityus. See Scorpions
of medical importance page for further information.
Unlike most other scorpion families, the Buthidae are of
both scientific and public interest due to their medical
importance in many parts of the world. Several species have
also made their way into the hobby. Many of these are safe to
be kept by experienced collectors, but the most dangerous
species should be avoided.
Androctonus crassicauda habitat picture from Kuwait are located HERE.
Photo of Parabuthus leiosoma (left)
by Jan Ove Rein (C) Photo of Hottentotta sp.
(right) by Jan Ove Rein (C)
SPECIES
FILES: This list of genera and
species is based on Fet
et al.(2000). Subspecies is not included in the list. I
try to update the list as additions and changes are published.
* denotes changes after Fet
et al. (2000). I will be grateful for information about
new development in the taxonomy of this family. For
information about synonyms and bibliographies, see Fet
et al. (2000) |
Afghanorthochirus
Lourenco & Vachon, 1997 A. daniellae
Lourenco & Vachon, 1997 A. erardi
Lourenco & Vachon, 1997 A. monodi
Lourenco & Vachon, 1997
|
Afroisometrus
Kovarík, 1997 A. minshullae (Fitzpatrick,
1994)
|
|
|
Alayotityus Armas,
1973
A. delacruzi Armas, 1973*
A. granma Armas, 1984 A.
juraguaensis Armas, 1973
A. lapidicola Teruel, 2002*
A. nanus
Armas, 1973
A. pallidus Teruel, 2002*
A. sierramaestrae Armas,
1973
|
Ananteris Thorell,
1891 A. ashmolei Lourenco, 1981 A.
balzanii Thorell, 1891 A.
charlescorfieldi Lourenco, 2001* A.
coinaui Lourenco, 1982 A. columbiana
Lourenco, 1991 A. cussinii Borelli,
1910 A. dekeyseri Lourenco, 1982 A.
deniseae Lourenco, 1997 A. ehrlichi
Lourenco, 1994 A. feae (Borelli,
1911) A. festae Borelli, 1899 A.
franckei Lourenco, 1982 A. gorgonae
Lourenco & Flórez, 1989 A. leilae
Lourenco, 1999* A. luciae Lourenco,
1984 A. maranhensis Lourenco, 1987 A.
mariaelenae Lourenco, 1999* A.
mariaterezae Lourenco, 1987 A. mauryi
Lourenco, 1982 A. platnicki Lourenco,
1993 A. pydanieli Lourenco, 1982 A.
sabineae Lourenco, 2001* A.
turumbanensis Gonzáles-Sponga, 1990 A.
venezuelensis Gonzáles-Sponga, 1972
|
|
|
Androctonus
Ehrenberg, 1828 A. amoreuxi (Audouin,
1826) A.
australis (Linnaeus, 1758) A.
bicolor Ehrenberg, 1828 A. crassicauda
(Olivier, 1807) A. finitimus (Pocock,
1897) A. hoggarensis (Pallary,
1929) A. mauritanicus (Pocock,
1902) A. sergenti Vachon, 1948
|
Anomalobuthus
Kraepelin, 1900 A. rickmersi Kraepelin,
1900
|
Apistobuthus
Finnegan, 1932 A. pterygocercus Finnegan,
1932 A. susanae Lourenco, 1998
|
Australobuthus
Locket, 1990 A. xerolimniorum Locket,
1990
|
Babycurus Karsch,
1886 B. ansorgei Hirst, 1911 B.
buettneri Karsch, 1886 B.
crassicaudatus Roewer, 1952 B.
crassimanus Caporiacco, 1936 B. gigas
Kraepelin, 1896 B. jacksoni (Pocock,
1890) B. johnstonii Pocock, 1896 B.
neglectus Kraepelin, 1897 B. ornatus
Werner, 1936 B. patrizii Borelli,
1925 B. somalicus Hirst, 1907 B.
subpunctatus Borelli, 1925 B.
taramassoi Borelli, 1919 B. wituensis
Kraepelin, 1913 B. zambonellii Borelli,
1902
|
Baloorthochirus
Kovarik, 1996 B. becvari Kovarík,
1996
|
Birulatus Vachon,
1974 B. haasi Vachon, 1974
|
Buthacus Birula,
1908 B. arenicola (Simon, 1885) B.
calviceps (Pocock, 1900) B. foleyi
Vachon, 1948 B. frontalis Werner,
1936
B. israelensis Lourenco, 2002*
B. leptochelys (Ehrenberg,
1829) B. spatzi (Birula, 1911) B.
villiersi Vachon, 1949 B. yotvatensis
Levy, Amitai & Shulov, 1973
|
Butheoloides
Hirst, 1925 B. annieae Lourenco,
1986 B. hirsti Lourenco, 1996 B.
maroccanus Hirst, 1925 B. milloti
Vachon, 1948 B. monodi Vachon,
1950 B. polisi Lourenco, 1996 B.
wilsoni Lourenco, 1995
|
Butheolus Simon,
1882 B. anthracinus (Pocock, 1895) B.
ferrugineus Kraepelin, 1898 B.
gallagheri Vachon, 1980 B. thalassinus
Simon, 1882
|
Buthiscus Birula,
1905 B. bicalcaratus Birula, 1905
|
Buthoscorpio
Werner, 1936 B. politus (Pocock,
1899) B. sarasinorum (Karsch,
1891)
|
Buthus Leach,
1815 B. atlantis Pocock, 1889 B.
barbouri Werner, 1932 B. insolitus
Borelli, 1925 B. maroccanus Birula,
1903 B. occitanus (Amoreux, 1789) |
Caribetityus
Lourenco, 1999* C. elii (Armas & Marcano
Fondeur, 1992)* C. quisqueyanus (Armas,
1982)* |
|
Centruroides Marx,
1890 C. alayoni Armas, 1999* C.
anchorellus Armas, 1976 C.
arctimanus (Armas, 1976) C. bani
Armas & Marcano Fondeur, 1987 C.
baracoae Armas, 1999* C. barbudensis
(Pocock, 1898) C. bertholdii (Thorell,
1876) C.
bicolor (Pocock, 1898) C.
chamulaensis Hoffmann, 1932 C.
chiapanensis Hoffmann, 1932 C. elegans
(Thorell, 1876) C.
excilicauda (Wood, 1863)
C. exilimanus Teruel & Stockwell, 2002*
C. exsul
(Meise, 1934) C. farri Armas, 1976 C.
flavopictus (Pocock, 1898) C. fulvipes
(Pocock, 1898) C.
gracilis (Latreille, 1804) C.
griseus (C. L. Koch, 1844) C.
guanensis Franganillo, 1930 C.
hentzi (Banks, 1900) C. hoffmanni
Armas, 1996 C. infamatus (C.L. Koch,
1844) C. insalanus (Thorell, 1876) C.
jaragua Armas, 1999* C. koesteri
Kraepelin, 1912 C.
limbatus (Pocock, 1898) C. limpidus
(Karsch, 1879) C. luceorum Armas,
1999* C. mahnerti Lourenco, 1983 C.
marcanoi Armas, 1981 C. margaritatus
(Gervais, 1841)
C. melanodactylus Teruel, 2002*
C. navarroi Teruel, 2002*
C. nigrescens (Pocock,
1898) C. nigrimanus (Pocock, 1898) C.
nigrovariatus (Pocock, 1898) C. nitidus
(Thorell, 1876) C. noxius Hoffman,
1932 C. ochraceus (Pocock, 1898) C.
pallidiceps Pocock, 1902 C. platnicki
Armas, 1981 C. pococki Sissom & Francke,
1983 C. rileyi Sissom, 1995 C.
robertoi Armas, 1976 C. schmidti
Sissom, 1995 C. sissomi Armas,
1996
C. stockwelli Teruel, 2002*
C. subgranosus (Kraepelin,
1898) C. suffusus (Pocock, 1902) C.
testaceus (DeGeer, 1778) C. thorellii
(Kraepelin, 1891) C. underwoodi Armas,
1976 C.
vittatus (Say, 1821)
|
Charmus Karsch,
1879 C. indicus Hirst, 1915 C.
laneus Karsch, 1879 C. sinhagadensis
Tikader & Bastawade, 1983
|
Cicileus Vachon,
1948 C. exilis (Pallary, 1928)
|
Compsobuthus
Vachon, 1949 C. abyssinicus (Birula,
1903) C. acutecarinatus (Simon,
1882) C. arabicus Levy, Amitai & Shulov,
1973 C. berlandi Vachon, 1950 C.
brevimanus (Werner, 1936) C. carmelitis
Levy, Amitai & Shulov, 1973 C. garyi
Lourenco & Vachon, 2001* C. jordanensis
Levy, Amitai & Shulov, 1973 C. klaptoczi
(Birula, 1909) C. longipalpis Levy, Amitai
& Shulov, 1973 C. maindroni (Kraepelin,
1900) C. manzonii (Borelli, 1915) C.
matthiesseni (Birula, 1905) C. polisi
Lowe, 2001* C. rugosulus (Pocock,
1900) C. vachoni Sisson, 1994 C.
werneri (Birula, 1908) |
Darchenia Vachon,
1977 D. bernadettae Vachon, 1977
|
Grosphus Simon,
1880 G. annulatus Fage, 1929 G.
bistriatus Kraepelin, 1900 G. feti
Lourenco, 1996 G. flavopiceus Kraepelin,
1900 G. grandidieri Kraepelin,
1900 G. hirtus Kraepelin, 1900 G.
limbatus (Pocock, 1889) G.
madagascariensis (Gervais, 1843)
|
Hemibuthus Pocock,
1900 H. crassimanus (Pocock, 1897)
|
Hemilychas Hirst,
1911 H. alexandrinus (Hirst, 1911)
|
Himalayotityobuthus
Lourenco, 1997 H. martensi Lourenco,
1997
|
Hottentotta
Birula, 1908 H. alticola (Pocock,
1895) H. arenaceus (Purcell, 1901) H.
conspersus (Thorell, 1876) H. eminii
(Pocock, 1890) H. franzwerneri (Birula,
1914) H. fuscitruncus (Caporiacco,
1936) H. hottentotta (Fabricius,
1787) H. jayakari (Pocock, 1895) H.
judaicus (Simon, 1872) H. kraepelini
(Roewer, 1943) H. minax (L. Koch,
1975) H. polystictus (Pocock,
1896) H. saulcyi (Simon, 1828) H.
scaber (Ehrenberg, 1828) H. schach
(Birula, 1905) H. socotrensis (Pocock,
1889) H. syrticus (Borelli, 1914) H.
trilineatus (Peters, 1861) H.
zagrosensis Kovarík, 1997
|
Iranobuthus
Kovarík, 1997 I. krali Kovarík,
1997
|
Isometroides
Keyserling, 1885 I. vescus (Karsch,
1880)
|
|
Isometrus Ehrenberg,
1828 I. acanthurus Pocock, 1899 I.
assamensis Oates, 1888 I. basilicus
Karsch, 1879 I. besucheti Vachon,
1982 I. brachycentrus Pocock, 1899 I.
corbeti Tikander & Batawade, 1983 I.
formosus Pocock, 1894 I. garyi
Lourenco & Huber, 2002* I. heimi Vachon,
1976 I. isadensis Tikander & Bastawade,
1983 I. krasenskyi Kovarík, 1998 I.
kurkai Kovarík, 1997 I. loebli
(Vachon, 1982)* I. maculatus
(DeGeer, 1778) I. melanodactylus (L. Koch,
1867) I. navaiae Kovarík, 1998 I.
rigidulus Pocock, 1897 I. sankeriensis
Tikander " Bastawade, 1983 I. thurstoni
Pocock, 1893 I. thwaitesi Pocock,
1897 I. vittatus Pocock, 1900 I.
zideki Kovarík, 1994
|
Karasbergia
Hewitt, 1913 K. methueni Hewitt, 1913 |
Kraepelinia
Vachon, 1974 K. palpator (Birula, 1903) |
Leiurus Ehrenberg,
1828
L. jordanensis Lourenco, Modry & Amr, 2002*
L.
quinquestriatus (Ehrenberg, 1828) |
Liobuthus Birula,
1898 I. kessleri Birula, 1898 |
Lissothus Vachon,
1948 L. bernardi Vachon, 1948 L.
occidentalis Vachon, 1950 |
Lychas C.L. Koch,
1845 L. albimanus Henderson, 1919 L.
asper (Pocock, 1891) L. biharensis
Tikader & Bastawade, 1983 L. braueri
(Kraepelin, 1986) L. buchardi Kovarík,
1997 L. burdoi (Simon, 1882) L.
farkasi Kovarik, 1997 L. feae (Thorell,
1889) L. flavimanus (Thorell,
1888) L. gravelyi Henderson, 1913 L.
hendersoni (Pocock, 1897) L.
heurtaultae Kovarík, 1997 L. hillyardi
Kovarík, 1997 L. hosei (Pocock,
1891) L. infuscatus (Pocock, 1891) L.
kamshetensis Tikader & Bastawade,
1983 L. kharpadi Bastawade, 1986 L.
krali Kovarík, 1995 L. laevifrons
Pocock, 1897 L. lourencoi Kovarík,
1997 L.
marmoreus (C.L. Koch, 1844) L.
mjobergi Kraepelin, 1916 L. mucronatus
(Fabricius. 1798) L. nigristernis (Pocock,
1899) L. obsti Kraepelin, 1913 L.
perfidus (Keyserling, 1885) L. rackae
Kovarík, 1997 L. rugosus (Pocock,
1897) L. scaber (Pocock, 1893) L.
scutilus C. L. Koch, 1845 L. serratus
(Pocock, 1891) L. shelfordi (Borelli,
1904) L. shoplandi (Oates, 1888) L.
srilankensis Lorenco, 1997 L.
tricarinatus (Simon, 1884) L. variatus
(Thorell, 1876) |
|
Lychasioides
Vachon, 1974 L. amieti Vachon, 1974 |
Mesobuthus Vachon,
1950 M. agnetis (Werner, 1936) M.
cyprius Gantenbein et al., 2000* M.
eupeus (C.L. Koch, 1839) M. extremus
(Werner, 1936) M. gibbosus (Brulli,
1832) M. hendersoni (Pocock, 1900) M.
macmahoni (Pocock, 1900) M. martensii
(Karsch, 1879) M. nigrocinctus (Ehrenberg,
1828)* M. pachyurus (Pocock, 1897) M.
rugiscutis (Pocock, 1897) M. tamulus
(Fabricius, 1798) M. vesiculatus (Pocock,
1899)
|
Mesotityus
Gonzalez-Sponga, 1981 M. vondangeli
González-Sponga, 1981 |
Microbuthus
Kraepelin, 1898 M. fagei Vachon,
1949 M. litoralis (Pavesi, 1885) M.
pusillus Kraepelin, 1898 |
Microtityus
Kjellesvig-Waering, 1966 M. biordi
González-Sponga, 1970 M. consuelo Armas
& Marcano Fondeur, 1987 M.
dominicanensis Santiago-Blay, 1985 M.
farleyi Teruel, 2000* M. fundorai
Armas, 1974 M. guantanamo Armas,
1984 M. iviei Armas, 1999* M.
jaumei Armas, 1974 M. joseantonioi
González-Sponga, 1981 M. lantiguai Armas
& Marcano Fondeur, 1992 M. paucidentatus
Armas & Marcano Fondeur, 1992 M. rickyi
Kjellesvig-Waering, 1966 M. trinitensis
Armas, 1974 M. vanzolinii Lorenco &
Eickstedt, 1983 M. virginiae Armas,
1999* M. waeringi Francke & Sissom, 1980
|
Nanobuthus Pocock,
1895 N. Andersoni Pocock, 1895 |
Neobuthus Hirst,
1911 N. berberensis (Hirst, 1911) |
Neogrosphus
Lourenco, 1995 N. blanci Lourenco,
1996 N. griveaudi (Vachon, 1969) |
Odontobuthus
Vachon, 1950 O. doriae (Thorell,
1876) O. odonturus (Pocock, 1897)
|
Odonturus Karsch,
1879 O.
dentatus Karsch, 1879 |
Olivierus
Farzanpay, 1987 O. caucasicus (Nordmann,
1840) |
Orthochiroides
Kovarík, 1998 O. vachoni Kovarík,
1998
|
Orthochirus
Karsch, 1891 O. aristidis (Simon,
1882) O. bicolor (Pocock, 1897) O.
flavescens (Pocock, 1897) O. fuscipes
(Pocock, 1900) O. innesi Simon,
1910 O. krishnai Tikader & Bastawade,
1983 O. luteipes Roewer, 1943 O.
pallidus (Pocock, 1897) O.
scrobiculosus (Grube, 1873)
|
Pakistanorthochirus
Lourenco, 1997 P. weitscahti Lourenco,
1997
|
Parabuthus Pocock,
1890 P. brevimanus (Thorell, 1876) P.
calvus Purcell, 1898 P. capensis
(Ehrenberg, 1831) P. distridor Lamoral,
1980 P. gracilis Lamoral, 1979 P.
granimanus Pocock, 1895 P.
granulatus (Ehrenberg, 1831) P.
heterurus Pocock, 1899 P. hunteri
Pocock, 1895 P. kalaharicus Lamoral,
1977 P. kraepelini Werner, 1902 P.
kuanyamarum Monrad, 1937 P. laevifrons
(Simon, 1888) P.
leiosoma (Ehrenberg, 1828) P.
maximus Werner, 1913 P. mixtus Borelli,
1925 P. mossambicensis (Peters,
1861) P. muelleri Prendini, 2000* P.
namibensis Lamoral, 1979 P. nanus
Lamoral, 1979 P. neglectus Purcell,
1899 P.
pallidus Pocock, 1895 P. planicauda
(Pocock, 1889) P. raudus (Simon,
1888) P. schlechteri Purcell, 1899 P.
stridulus Hewitt, 1913 P.
transvaalicus Purcell, 1899 P.
villosus (Peters, 1862) P. zavattarii
Caporiacco, 1939
|
Paraorthochirus
Lourenco & Vachon, 1997
P. blandini Lourenco & Vachon, 1997
P. glabrifrons
(Kraepelin, 1903) P. goyffoni Lourenco &
Vachon, 1995
P. kaspareki Lourenco & Huber, 2000*
P. kinzelbachi Lourenco & Huber, 2000*
P. stockwelli Lourenco &
Vachon, 1995 |
Pectinibuthus Fet,
1984 P. birulai Fet, 1984 |
Plesiobuthus
Pocock, 1900 P. paradoxus Pocock, 1900
|
Polisius Fet,
Capes & Sissom, 2001* P. persicus Fet,
Capes & Sissom, 2001* |
|
Psammobuthus
Birula, 1911 P. zarudnyi Birula, 1911
|
Pseudolissothus
Lourenco, 2001* P. pussilus Lourenco, 2001*
|
Pseudolychas
Kraepelin, 1911 P. ochraceus (Hirst,
1911) P. pegleri (Purcell, 1901) P.
transvaalicus Lawrence, 1961
|
|
Pseudouroplectes
Lourenco, 1995 P. betschi Lourenco, 1995 |
Razianus
Farzanpay, 1987 R. zarudnyi (Birula, 1903)
|
Rhopalurus
Thorell, 1876 R. abudi Armas & Marcano
Fondeur, 1987 R. acromelas Lutz & Mello,
1922 R. agamemmon (C. L. Koch,
1839) R. amazonicus Lourenco, 1986 R.
debilis (C. L. Koch, 1840) R.
garridoi Armas, 1974 R.
junceus (Herbst, 1800) R. lacrau
Lourenco & Pinto-da-Rocha, 1997 R.
laticauda Thorell, 1876 R. piceus
Lourenco & Pinto-da-Rocha, 1997 R.
princeps (Karsch, 1879) R. rochai
Borelli, 1910 |
Sassandiothus
Farzanpay, 1987 S. zarudnyi (Birula, 1900)
|
Simonoides Vachon
& Farzanpay, 1987 S. farzanpayi Vachon
& Farzanpay, 1987 |
Somalibuthus
Kovarík, 1998 S. demisi Kovarík, 1998 |
Somalicharmus
Kovarík, 1998 S. withmanae Kovarík, 1998 |
Thaicharmus
Kovarík, 1995 T. mahunkai Kovarík, 1995
|
Tityobuthus
Pocock, 1893 T. baroni (Pocock,
1890)
T. darainensis Lourenco & Goodman, 2002*
T. dastychi Lourenco, 1997 T.
gracilis (Fage, 1946) T. guillaumeti
Lourenco, 1995 T. judsoni Lourenco,
1996 T. lucileae Lourenco, 1996 T.
parrilloi Lourenco, 1996 T. petrae
Lourenco, 1996 T. pococki Lourenco, 1995 |
Tityopsis Armas,
1974 T. inaequalis (Armas, 1974) T.
inexpectata (Moreno, 1940) |
Tityus C.L. Koch,
1836 T. abudi Armas, 1999* T.
acutidens Mello-Leităo, 1933 T.
altithronus Armas, 1999* T.
androcottoides (Karsch, 1879) T.
anduzei González-Sponga, 1997 T. anneae
Lourenco, 1997 T. antioquensis Lourenco
& Otero Patińo, 1998
T. apiacas Lourenco 2002*
T. arellanoparrai
González-Sponga, 1985 T. argentinus Borelli,
1899 T. asthenes Pocock, 1893 T.
atriventer Pocock, 1897 T. bahiensis
(Perty, 1833) T. barquisimetanus
González-Sponga, 1994 T. bastosi Lourenco,
1984 T. bellulus Armas, 1999* T.
betschi Lourenco, 1992 T. birabeni
Abalos, 1955 T. blanci Lourenco,
1994 T. blaseri Mello-Leităo, 1931 T.
boconoensis González-Sponga, 1981 T.
bolivanus Kraepelin, 1895 T. brazilae
Lourenco & Eickstedt, 1984 T. breweri
González-Sponga, 1997 T. carabobensis
González-Sponga, 1987 T. carinatoides
Mello-Leităo, 1945 T. carvalhoi
Mello-Leităo, 1945 T. cerroazul Lourenco,
1986 T. charalaensis Mello-Leităo,
1940 T. charreyroni Vellard, 1932 T.
clathratus C. L. Koch, 1844 T.
columbianus (Thorell, 1876) T.
confluens Borelli, 1899 T. costatus
(Karsch, 1879) T. crassimanus (Thorell,
1876) T. cuellari Lourenco, 1994 T.
culebrensis González-Sponga, 1994 T.
dasyurus Pocock, 1897 T. dedoslargos
Francke & Stockwell, 1987 T. demangei
Lourenco, 1981 T. dinizi Lourenco,
1997 T. discrepans (Karsch, 1879) T.
dupouyi González-Sponga, 1987 T.
ebanoverde Armas, 1999* T. ecuadorensis
Kraepelin, 1896 T. engelkei Pocock,
1902 T. erikae Lourenco, 1999* T.
evandroi Mello-Leităo, 1945 T.
exstinctus Lourenco, 1995 T.
falconensis González-Sponga, 1974 T.
fasciolatus Pessôa, 1935 T. festae
Borelli, 1899 T. filodendron
González-Sponga, 1981 T. florezi Lourenco,
2000* T. footei Chamberlin, 1916 T.
forcipula (Gervais, 1843) T. fuehrmanni
Kraepelin, 1914 T. funestus Hirst,
1911 T. gaffini Lourenco, 2000* T.
gasci Lourenco, 1982 T. indecisus
Mello-Leităo, 1934 T. insignis (Pocock,
1889) T. intermedius Borelli, 1899 T.
isabelceciliae González-Sponga, D'Suze &
Sevcik, 2001* T. ivicnancor González-Sponga,
1997
T. jeanvellardi Lourenco, 2001*
T. jussarae Lourenco, 1988 T.
kuryi Lourenco, 1997 T. lamottei
Lourenco, 1981 T. lancinii González-Sponga,
1972 T. lourencoi Flórez, 1996 T.
lutzi Giltay, 1928 T. macrochirus
Pocock, 1897
T. magnimanus Pocock,
1897
T. mattogrossensis Borelli,
1901
T. melanostictus Pocock, 1893
T. meridanus González-Sponga, 1981 T.
metuendus Pocock, 1897 T. michelii
Armas, 1982 T. microcystis Lutz & Mello,
1922 T. monaguensis González-Sponga,
1974 T. mongei Lourenco, 1996 T.
munozi Lourenco, 1997 T. neglectus
Mello-Leităo, 1932 T. neibae Armas,
1999* T. nematochirus Mello-Leităo,
1940 T. neoespartanus González-Sponga,
1996 T. nororientalis González-Sponga,
1996 T. obtusus (Karsch, 1879) T.
ocelote Francke & Stockwell, 1987 T.
osmanus González-Sponga, 1996 T. oteroi
Lourenco, 1998 T. ottenwalderi Armas,
1999* T. pachyurus Pocock, 1897 T.
paraensis Kraepelin, 1896 T.
paraguayensis Kraepelin, 1895 T.
parvulus Kraepelin, 1914 T.
perijanensis González-Sponga, 1994 T.
pictus Pocock, 1893 T. pittieri
González-Sponga, 1981 T. pococki Hirst,
1907 T. portoplatensis Armas & Marcano
Fondeur, 1992 T. prancei Lourenco,
2000* T. proseni Abalos, 1954 T.
pugilator Pocock, 1898 T. pusillus
Pocock, 1893 T. ramirezi Esquivel de Verde,
1969 T. raquelae Lourenco, 1988 T.
rebieri Lourenco, 1997 T. riocaurensis
González-Sponga, 1996 T. roigi Maury &
Lourenco, 1987 T. rojasi González-Sponga,
1996 T. rufofuscus Pocock, 1897 T.
rugosus Schenkel, 1932 T. rusmelyae
González-Sponga, D'Suze & Sevcik, 2001* T.
sabinae Lourenco, 1994 T. sanarensis
González-Sponga, 1997 T. sastrei Lourenco
& Flórez, 1990 T. serrulatus Lutz &
Mello, 1922 T. shiriana González-Sponga,
1991 T. silvestris Pocock, 1897 T.
simonsi Pocock, 1900 T. soratensis
Kraepelin, 1912 T. stigmurus (Thorell,
1876) T. strandi Werner, 1939 T.
surorientalis González-Sponga, 1996 T.
tamayoi González-Sponga, 1987 T.
tayrona Lourenco, 1991 T. thelyacanthus
Mello-Leităo, 1933 T. trinitatis Pocock,
1897 T. trivittatus Kraepelin,
1898 T. tucurui Lourenco, 1988 T.
uniformis Mello-Leităo, 1931 T. urbinai
Scorza, 1952
T. uruguayensis Borelli,
1901
T. vaissadei Lourenco, 2002*
T. valerae Scorza, 1954 T.
venamensis González-Sponga, 1981 T.
zulianus González-Sponga, 1981
|
Uroplectes Peters,
1861 U. fischeri (Karsch, 1879) U.
andreae Pocock, 1889 U. carinatus
(Pocock, 1890) U. chubbi Hirst,
1911 U. fischeri (Karsch, 1879) U.
flavoviridis Peters, 1861 U. formosus
Pocock, 1890 U. gracilior Hewitt,
1914 U. insignis Pocock, 1890 U.
lineatus (C. L. Koch, 1844) U.
longimanus Werner, 1936 U. marlothi
Purcell, 1901 U. ngangelarum Monard,
1930 U. occidentalis Simon, 1876 U.
olivaceus Pocock, 1896 U.
otjimbinguensis (Karsch, 1879) U.
pardalis Werner, 1913 U. patrizii
Caporiacco, 1936 U. pictus Werner,
1913 U. pilosus (Thorell, 1876) U.
planimanus (Karsch, 1879) U.
schlechteri Purcell, 1901 U. schubotzi
Kraepelin, 1929 U. silvestrii Borelli,
1913 U. teretipes Lawrence, 1966 U.
triangulifer (Thorell, 1876) U.
tumidimanus Lamoral, 1979 U. variegatus
(C. L. Koch, 1844) U. vittatus (Thorell,
1876) U. xanthogrammus Pocock,
1897
|
Uroplectoides
Lourenco, 1998 U. abyssinicus Lourenco,
1998 U. emiliae (Werner, 1916) |
Vachoniolus Levi,
Amitai & Shulov, 1973 V. globimanus
Levy, Amitai & Shulov, 1973 V.
minipectinatus (Levy, Amitai & Shulov, 1973)
|
Vachonus Tikader
& Bastawade, 1983 V. astrostriatus
(Pocock, 1897) V. rajasthanicus Tikader
& Bastawade, 1983
|
Zabius Thorell,
1893 Z. birabeni Mello-Laeităo,
1938 Z. fuscus (Thorell, 1876)
| | |